A womanizing city man meets Tillie in the country. When he sees that her father has a very large bankroll for his workers, he persuades her to elope with him.

Tillie Todd gifts her wealthy Aunt Sally a dime-store tomato pincushion, falsely claiming it’s handmade. Cousin Percy exposes the lie, vying for Sally’s favor. At Sally’s birthday, Tillie’s suitor “The Bat” crash-lands his flying invention, inspiring Sally to take flight herself. Declared dead after vanishing for days, she wills Tillie the pincushion but leaves most of her estate to her pet monkey Jim, with Percy as eventual heir. Percy kidnaps Jim to hasten his fortune, framing Tillie for the crime. Pursued by police, Tillie is saved when Sally returns alive. In frustration, Tillie tears apart the despised pincushion—only to discover it stuffed with $10,000.

Tillie and her neighbor Mr. Pipkins are both distraught over their respective marriages. One day, they sneak off to have a lively time at Coney Island. They flee the park together just as their spouses come to find them. After a chase, each is rescued from the ocean and reconcile with their respective spouses.